• rapid diversification of multicellular animal life ~541 million years ago
  • most major animal phyla appeared within a ~25 million year window
  • origin of eyes, shells, exoskeletons, predation, bilateral symmetry
  • preceded by the Ediacaran biota: soft-bodied, sessile organisms
  • triggers: rising oxygen levels, calcium availability, ecological arms races
  • first appearance of arthropods, mollusks, chordates, echinoderms
  • the Burgess Shale and Chengjiang fossil beds preserve this radiation
  • deepest example of an epoch boundary driven by innovation cascades
  • analogous to the Information Age explosion of digital complexity